Documentation for JIRA 4.3. Documentation for other versions of JIRA is available too. 
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JIRA 4.3 and later adds classes to the HTML tag on page load. This means that you can now put all browser-specific CSS fixes in the main stylesheet near related styles. Having styles in one file increases maintainability, and reduces the number of requests — which helps pages load faster.
Prior to 4.3 you would put IE styles in the IE stylesheet and prefix with * or _ depending on the browser. In JIRA 4.3 and later you can use the following: